Mandibular molar

The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name molar derives from Latin, molaris dens, meaning 'millstone tooth', from mola, millstone and dens, tooth. Molars show a great deal of diversity in size and shape across mammal groups. The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name molar derives from Latin, molaris dens, meaning 'millstone tooth', from mola, millstone and dens, tooth. Molars show a great deal of diversity in size and shape across mammal groups. In humans, the molar teeth have either four or five cusps. Adult humans have 12 molars, in four groups of three at the back of the mouth. The third, rearmost molar in each group is called a wisdom tooth. It is the last tooth to appear, breaking through the front of the gum at about the age of 20, although this varies from individual to individual. Race can also affect the age at which this occurs, with statistical variations between groups. In some cases, it may not even erupt at all. The human mouth contains upper (maxillary) and lower (mandibular) molars. They are: maxillary first molar, maxillary second molar, maxillary third molar, mandibular first molar, mandibular second molar, and mandibular third molar. In mammals, the crown of the molars and premolars is folded into a wide range of complex shapes. The basic elements of the crown are the more or less conical projections called cusps and the valleys that separate them. The cusps contain both dentine and enamel, whereas minor projections on the crown, called crenullations, are the result of different enamel thickness. Cusps are occasionally joined to form ridges and expanded to form crests. Cingula are often incomplete ridges that pass around the base of the crown. Mammalian, multicusped cheek teeth probably evolved from single-cusped teeth in reptilians, although the diversity of therapsid molar patterns and the complexity in the molars of the earliest mammals make determining how this happened impossible. According to the widely accepted 'differentiation theory', additional cusps have arisen by budding or outgrowth from the crown, while the rivalling 'concrescence theory' instead proposes that complex teeth evolved by the clustering of originally separate conical teeth. Therian mammals (placentals and marsupials) are generally agreed to have evolved from an ancestor with tribosphenic cheek teeth, with three main cusps arranged in a triangle. Each major cusp on an upper molar is called a cone and is identified by a prefix dependent on its relative location on the tooth: proto-, para-, meta-, hypo-, and ento-. Suffixes are added to these names: -id is added to cusps on a lower molar (e.g., protoconid); -ule to a minor cusp (e.g., protoconulid). A shelf-like ridge on the lower part of the crown (on an upper molar) is called a cingulum; the same feature on the lower molar a cingulid, and a minor cusp on these, for example, a cingular cuspule or conulid. The design that is considered one of the most important characteristics of mammals is a three-cusped shape called a tribosphenic molar. This molar design has two important features: the trigonid, or shearing end, and the talonid, or crushing heel. In modern tribosphenic molars, the trigonid is towards the front of the jaw and the talonid is towards the rear. The tribosphenic tooth is found in insectivores and young platypuses (adults have no teeth). Upper molars look like three-pointed mountain ranges; lowers look like two peaks and a third off to the side. The tribosphenic design appears primitively in all groups of mammals. Some paleontologists believe that it developed independently in monotremes (or australosphenidans), rather than being inherited from an ancestor that they share with marsupials and placentals (or boreosphenidans); but this idea has critics and the debate is still going on. For example, the dentition of the Early Cretaceous monotreme Steropodon is similar to those of Peramus and dryolestoids, which suggests that monotremes are related to some pre-tribosphenic therian mammals, but, on the other hand, the status of neither of these two groups is well-established.